Rachel Whetstone, the Briton who has been head of communications and public policy at Google for several years, has been poached by Uber for a similar role.
She will become senior vice-president of policy and communications at the taxi company, which is expected to list on the stock market this year and has been valued at as much as $50bn (£31.7bn).
Whetstone will replace David Plouffe, who was Barack Obama’s campaign manager in 2008 and joined Uber in August last year.
He has been elevated to the role of chief adviser to the company and its chief executive Travis Kalanick, and will also sit on the board of Uber.
Whetstone’s appointment was first reported by Recode. She begins her new job next month.
In recent months Uber has been buffeted on several fronts in a number of cities and countries by regulators, taxi drivers and others who oppose the rise of the smartphone-based service that has proved popular with thousands of users.
In the face of fierce resistance in numerous European cities, Kalanick said in January that he wanted to focus on forging “new partnerships”. Uber’s expansion could eventually create 50,000 jobs, he added.
Whetstone is married to Steve Hilton, who was a senior adviser to David Cameron before quitting in 2012. He now lives in California and is chief executive of technology start-up Crowdpac.
Policy Exchange, a conservative British thinktank, said Hilton would become a visiting scholar and visit the UK several times a year but not be based in London.
Before starting her career with Google in Europe in 2005, Whetstone was Michael Howard’s chief of staff after he became Conservative leader in 2003.
She joined Conservative Central Office after studying politics at Bristol University before switching to Carlton Communications – where she worked with David Cameron – and then went on to co-found the PR agency Portland.
The couple were godparents to Ivan, the Camerons’ disabled son who died in 2009.
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